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(*denotes student or postdoc from Ji lab)


*Iles, D. T., H. Lynch, R. Ji, C. Barbraud, K. Delord, and S. Jenouvrier. 2020. Sea ice predicts long-term trends in Adélie penguin population growth, but not annual fluctuations: Results from a range-wide multiscale analysis. Global Change Biology, 26(7):3788–3798,

Jenouvrier, S., M. Holland, D. Iles, S. Labrousse, L. Landrum, J. Garnier, H. Caswell, H. Weimerskirch, M. LaRue, R. Ji, and C. Barbraud. 2020. The Paris Agreement objectives will likely halt future declines of emperor penguins. Global Change Biology, 26(3):1170–1184,



Ji, R., M. Jin, Y. Li, Y.-H. Kang, and C.-K. Kang. 2019. Variability of primary production among basins in the East/Japan Sea: Role of water column stability in modulating nutrient and light availability. Progress in Oceanography, 178:102173.P-60.

*Kvile, K. Ø., C. Ashjian, and R. Ji. 2019. Pan-Arctic depth distribution of diapausing Calanus copepods. The Biological Bulletin, 237(2):76-89,

Bandara, K., Ø. Varpe, R. Ji, and K. Eiane. 2019. Artificial evolution of behavioral and life history strategies of high-latitude copepods in response to bottom-up and top-down selection pressures. Progress in Oceanography, 173:134–164.

Ji, B. Y., Z. O. Sandwith, W. J. Williams, O. Diaconescu, R. Ji, Y. Li, E. V. Scoy, M. Yamamoto‐Kawai, S. Zimmermann, and R. H. R. Stanley. 2019. Variations in Rates of Biological Production in the Beaufort Gyre as the Arctic Changes: Rates From 2011 to 2016. Journal of Geophysical Research: Oceans, 124(6):3628–3644.

Labrousse, S., A. D. Fraser, M. Sumner, T. Tamura, D. Pinaud, B. Wienecke, R. Kirkwood, Y. Ropert‐Coudert, R. Reisinger, I. Jonsen, R. Porter‐Smith, C. Barbraud, C.-A. Bost, R. Ji, and S. Jenouvrier. 2019. Dynamic Fine-Scale Sea Icescape Shapes Adult Emperor Penguin Foraging Habitat in East Antarctica. Geophysical Research Letters, 46(20):11206–11218,

Record, N, J. Runge, D. Pendleton, W. Balch, K. Davies, A. Pershing, C. Johnson, K. Stamieszkin, R. Ji, Z. Feng, S. Kraus, R. Kenney, C. Hudak, C. Mayo, C. Chen, J. Salisbury, and C. Thompson. 2019. Rapid Climate-Driven Circulation Changes Threaten Conservation of Endangered North Atlantic Right Whales. Oceanography, 32(2)

Staudinger, M. D., K. E. Mills, K. Stamieszkin, N. R. Record, C. A. Hudak, A. Allyn, A. Diamond, K. D. Friedland, W. Golet, M. E. Henderson, C. M. Hernandez, T. G. Huntington, R. Ji, C. L. Johnson, D. S. Johnson, A. Jordaan, J. Kocik, Y. Li, M. Liebman, O. C. Nichols, D. Pendleton, R. A. Richards, T. Robben, A. C. Thomas, H. J. Walsh, and K. Yakola. 2019. It’s about time: A synthesis of changing phenology in the Gulf of Maine ecosystem. Fisheries Oceanography,



*Kvile, Ø. K., C. J. Ashjian, Z. Feng, J. Zhang, R. Ji, 2018. Pushing the limit: Resilience of an Arctic copepod to environmental fluctuations. Global Change Biology. DOI: 10.1111/gcb.14419.

Record, N. R.  R. Ji, F. Maps, Ø. Varpe, J. A. Runge, C. M. Petrik, D. Johns, 2018. Copepod diapause and the biogeography of the marine lipidscape. Journal of Biogeography. DOI:

Xu, T., J. Sun, H.K. Watanabe, C. Chen, M. Nakamura, R. Ji, D. Feng, J. Lv, S. Wang, Z. Bao, P.Y. Qian, J.W. Qiu, 2018. Population connectivity, genetic structure and environmental adaptation of deep-sea mussels as revealed by genome-wide SNP markers. Molecular Ecology.  DOI: 10.1111/eva.12696.

*Feng, Z., R. Ji, C. Ashjian, R. Campbell, J. Zhang, 2018. Biogeographic responses of the copepod Calanus glacialis to a changing Arctic marine environment. Global. Change Biology 24:e159–e170. doi:10.1111/gcb.13890.

Maar, M., M. Butenschön, U. Daewel, A. Eggert, W. Fan, S. S. Hjøllo, M. Hufnagl, M. Huret, R. Ji, G. Lacroix, M. A. Peck, H. Radtke, S. Sailley, M. Sinerchia, M. D. Skogen, M. Travers-Trolet, T. A. Troost, and K. van de Wolfshaar. 2018. Responses of summer phytoplankton biomass to changes in top-down forcing: Insights from comparative modelling. Ecological Modelling, 376:54–67.

Bandara, K., Ø. Varpe, R. Ji, and K. Eiane, 2018. A high-resolution modeling study on diel and seasonal vertical migrations of high-latitude copepods. Ecological Modelling 368: 357–376. doi:10.1016/j.ecolmodel.2017.12.010.



Ji, R., Z. Feng, B. T. Jones, C. Thompson, C. Chen, N. R. Record, and J. A. Runge, 2017. Coastal amplification of supply and transport (CAST): a new hypothesis about the persistence of Calanus finmarchicus in the Gulf of Maine. ICES J Mar Sci. doi:10.1093/icesjms/fsw253.

Youngflesh, C., Jenouvrier, S., Li, Y., Ji, R., Ainley, D.G., Ballard, G., Barbraud, C., Delord, K., Dugger, K.M., Emmerson, L.M., Fraser, W.R., Hinke, J.T., Lyver, P.O., Olmastroni, S., Southwell, C.J., Trivelpiece, S.G., Trivelpiece, W.Z., Lynch, H.J., 2017. Circumpolar analysis of the Adélie Penguin reveals the importance of environmental variability in phenological mismatch. Ecology 98, 940–951. doi:10.1002/ecy.1749

Payne, M.R., Hobday, A.J., MacKenzie, B.R., Tommasi, D., Dempsey, D.P., Fässler, S.M.M., Haynie, A.C., Ji, R., Liu, G., Lynch, P.D., Matei, D., Miesner, A.K., Mills, K.E., Strand, K.O., Villarino, E., 2017. Lessons from the First Generation of Marine Ecological Forecast Products. Front. Mar. Sci. 4. doi:10.3389/fmars.2017.00289

Che-Castaldo, C, S Jenouvrier, C Youngflesh, K Shoemaker, G Humphries, P McDowall, L Landrum, M Holland, Y Li, R Ji, HJ Lynch. 2017. Spatial aggregation reveals robust dynamics despite stochastic noise in pan-Antarctic analysis of Adélie penguin abundance. Nature Communications, doi:10.1038/s41467-017-00890-0



*Feng, Z., R. Ji, R. G. Campbell, C. J. Ashjian, and J. Zhang. 2016. Early ice retreat and ocean warming may induce copepod biogeographic boundary shifts in the Arctic Ocean. J. Geophys. Res. Oceans. doi:10.1002/2016JC011784

*Li, Y., R. Ji, S. Jenouvrier, M. Jin, and J. Stroeve, 2016. Synchronicity between ice retreat and phytoplankton bloom in circum-Antarctic polynyas, Geophysical Research Letters, doi: 10.1002/2016gl067937.

*Jones, BT and Solow, A and Ji, R, 2016. Resource allocation for Lagrangian tracking. Journal of Atmospheric and Oceanic Technology.

Jin, M., Popova, E. E., Zhang, J., Ji, R., Pendleton, D., Varpe, Ø., … Lee, Y. J. 2016. Ecosystem model intercomparison of under-ice and total primary production in the Arctic Ocean. Journal of Geophysical Research: Oceans, 121(1), 934–948.

Friedland, K. D., Record, N. R., Asch, R. G., Kristiansen, T., Saba, V. S., Drinkwater, K. F., … Ji, R. 2016. Seasonal phytoplankton blooms in the North Atlantic linked to the overwintering strategies of copepods. Elementa: Science of the Anthropocene, 4, 000099.



Sathyendranath, S., R. Ji, and H. I. Browman, 2015. Introduction to the Themed Section: ‘Revisiting Sverdrup’s Critical Depth Hypothesis’.  ICES Journal of Marine Science, 72(6), 1892–1896. doi:10.1093/icesjms/fsv110.  (Editor for the special issue).

*Li, Y., P. S. Fratantoni, C. Chen, J. A. Hare, Y. Sun, andR. C. Beardsley, R. Ji, 2015. Spatio-temporal patterns of stratification on the Northwest Atlantic shelf, Progress in Oceanography, doi:10.1016/j.pocean.2015.01.003



Friedland, K. D., R. T. Leaf, J. Kane, D. Tommasi, R. G. Asch, N. Rebuck, R. Ji, S. I. Large, C. Stock, V. S. Saba, 2014. Spring Bloom Dynamics and Zooplankton Biomass Response on the US Northeast Continental Shelf. Continental Shelf Research 102:47-61

Llopiz, J.K., R.K. Cowen, M.J. Hauff, R. Ji, P.L. Munday, B.A. Muhling, M.A. Peck, D.E. Richardson, S. Sogard, and S. Sponaugle. 2014. Early life history and fisheries oceanography: New questions in a changing world. Oceanography 27(4):26–41,

Petrik, C.M.  R. Ji, C. Davis, 2014.  Interannual differences in larval haddock survival: hypothesis testing with a 3D biophysical model of Georges Bank.   Fisheries Oceanography, 23(6): 521-553.

Runge, J.A., R. Ji, C. Thompson, N. Record, F. Maps, C. Chen, J. Salisbury, D. Vandemark, 2014. Persistence of Calanus finmarchicus in the western Gulf of Maine under strong environmental forcing. Journal of Plankton Research, DOI: 10.1093/plankt/fbu098.

*Li, Y., R. Ji, P. S. Fratantoni, C. Chen, J. A. Hare, C. S. Davis, and R. C. Beardsley (2014), Wind-induced interannual variability of sea level slope, along-shelf flow, and surface salinity on the Northwest Atlantic shelf, J. Geophys. Res. Oceans, doi:10.1002/2013JC009385.

Tian, R., C. Chen, J. Qi, R. Ji, R. C. Beardsley and C. S. Davis, 2014. Model Study of Nutrient and Phytoplankton Dynamics in the Gulf of Maine: Patterns and Drivers for Seasonal and Interannual Variability.  ICES Journal of Marine Sciences, 10.1093/icesjms/fsu090. 72(2): 388-402

Bi, H., R. Ji, H. Liu, Y.-H. Jo, and J. A. Hare, 2014. Decadal Changes in Zooplankton of the Northeast U.S. Continental Shelf, PLoS ONE, 9(1), e87720, doi:10.1371/journal.pone.0087720.

Li, Y., X. Chen, C. Chen, J. Ge, R. Ji, R. Tian, P. Xue, and L. Xu, 2014. Dispersal and survival of chub mackerel (Scomber Japonicus) larvae in the East China Sea, Ecological Modelling, 283, 70–84, doi:10.1016/j.ecolmodel.2014.03.016.

Chen, C., Z. Lai, R. C. Beardsley, J. Sasaki, J. Lin, H. Lin, R. Ji, and Y. Sun, 2014. The March 11, 2011 Tōhoku M9.0 earthquake-induced tsunami and coastal inundation along the Japanese coast: A model assessment, Progress in Oceanography, 123, 84–104, doi:10.1016/j.pocean.2014.01.002.



Ji, R., M. Jin, Ø. Varpe, 2013. Sea ice phenology and timing of primary production pulses in the Arctic Ocean. Global Change Biology, DOI: 10.1111/gcb.12074. PDF

Batchelder, H., K. Daly, C. Davis, R. Ji, M. Ohman, W. Peterson, and J. Runge, 2013. Climate Impacts on Zooplankton Population Dynamics in Coastal Marine Ecosystems, Oceanography, 26(4), 34–51, doi:10.5670/oceanog.2013.74.

Lai, Z., C. Chen, R.C. Beardsley, H. Lin, R. Ji, J. Sasaki and J. Lin, 2013. Initial spread of 137Cs over the shelf of Japan: a study using the high-resolution global-coastal nesting ocean model, Biogeosciences. 10, 5439–5449.

Daewel, D., S. Hjøllo,M. Huret,R. Ji, M. Maar, S, Niiranen, M. Travers-Trolet, M. Peck, K. van de Wolfshaar, 2013. Trophic control of zooplankton dynamics: a review on observations and models.  ICES Journal of Marine Science. 10.1093/icesjms/fst125.

Greene, C., E. Meyer-Gutbrod, B. Monger, L. McGarry, A. Pershing, I. Belkin, P. Fratantoni, D. Mountain, R. Pickart, A. Proshutinsky, R. Ji, J. Bisagni, S. Hakkinen, D. Haidvogel, J. Wang, E. Head, P. Smith, C. Reid, A. Conversi, 2013. Remote climate forcing of decadal-scale regime shifts in Northwest Atlantic shelf ecosystems.  Limnology and Oceanography, 58(3), 2013, 803-816.



Ji, R., *C. Stegert, C. Davis, 2012. Sensitivity of copepod populations to bottom-up and top-down forcing: a modeling study in the Gulf of Maine region. Journal of Plankton Research, doi: 10.1093/plankt/fbs070. PDF

Ji, R., C. Ashjian, R. Campbell, C. Chen, G. Gao, C. Davis, G. Cowles, R. Beardsley, 2012. Life history and biogeography of Calanus copepods in the Arctic Ocean: An individual-based modeling study.  Progress in Oceanography. 96:40-56. PDF

*Stegert, C. R. Ji, N. Li, C. Davis, 2012. Processes controlling seasonality and spatial distribution of Centropages typicus: A modeling study in the Gulf of Maine/Georges Bank region. Journal of Plankton Research. 34(1): 18-35. PDF

Greene C. H., coauthros, R. Ji, coauthors, 2012. Remote Climate Forcing of Regime Shifts in Northwest Atlantic Shelf.  Oceanography. 25(3):208–213, PDF

*Song, X.  Z. Lai, R. Ji, C. Chen, L. Huang, J. Yin, Y. Wang, S. Lian, X. Zhu, 2012. Summertime primary production in northwest South China Sea: Interaction of coastal eddy, upwelling and biological processes.   Continental Shelf Research, doi:10.1016/j.csr.2012.07.016



Ji, R. 2011.  Calanus finmarchicus diapause initiation: new view from traditional life history based model.  Marine Ecology Progress Series440:105-114. PDF

*Song, H., R. Ji, C. Stock, K. Kearney, Z. Wang, 2011. Interannual variability of phytoplankton bloom and plankton productivity in the Nova Scotian Shelf and Gulf of Maine. Marine Ecology Progress Series. 426:105-118. PDF

Hu, S., C. Chen, Ji, R., et al., 2011.  Impact of physical forcings on the interannual variability of fall phytoplankton bloom: a mixed-layer box model and a 1-D numerical model studies.  Marine Ecology Progress Series. 427: 29-49. PDF 



 Ji, R., M. Edwards, D.L. Mackas, J. Runge, A.C. Thomas, 2010. Marine plankton phenology and life history in a changing climate:Current research and future directions.  Journal of Plankton Research - Horizon, 32(10): 1355–1368, doi:10.1093/plankt/fbq062. PDF.

*Stegert, C., Ji, R., Davis, C., 2010. Influence of ocean warming on population growth rates in North Altantic copepod species: An empirical projection. Progress in Oceanography. 87:264-276. PDF

Runge J., coauthers, R. Ji, coauthors, 2010. Understanding Climate Impacts on Recruitment and Spatial Dynamics of Atlantic Cod in the Gulf of Maine: Integration of Observations and Modeling. Progress in Oceanography. doi:10.1016/j.pocean.2010.09.016. PDF

*Song, H., R. Ji, C. Stock, Z. Wang, 2010. Phenology of phytoplankton blooms in the Nova Scotian Shelf–Gulf of Maine region: remote sensing and modeling analysis.Journal Plankton Research. doi:10.1093/plankt/fbq086. PDF.

*Lin, P., R. Ji, C. Davis, D. McGillicuddy, 2010. Optimizing plankton survey strategies using Observing System Simulation Experiments.  Journal of Marine Systems, 82: 187-194. PDF


Before 2010

Ji, R., C. Davis, C. Chen, R. Beardsley, 2009.  Life history traits and spatiotemporal distributional patterns of copepod populations in the Gulf of Maine-Georges Bank region.  Marine Ecology Progress Series, 384:187-205. PDF

Ji, R., C. Davis, C. Chen, D. Townsend, D. Mountain, R. Beardsley, 2008.  Modeling the influence of low-salinity water inflow on winter-spring phytoplankton dynamics in the Nova Scotian Shelf – Gulf of Maine region. Journal of Plankton Research, 30(12):1399-1416. PDF

Ji, R., C. Davis, C. Chen, and R. Beardsley, 2008. Influence of local and external processes on the annual nitrogen cycle and primary productivity on Georges Bank: A 3-D biological-physical modeling study. Journal of Marine Systems, 73:31-47. PDF

Hu, S., D. Townsend, C. Chen, G. Cowles, R. Beardsley, R. Ji, R. Houghton, 2008. Tidal pumping and nutrient fluxes on Georges Bank: A process-oriented modeling study. Journal of Marine Systems, 74: 528-544. PDF

Ji, R., C. Davis, C. Chen, D. Townsend, D. Mountain, R. Beardsley, 2007.  Influence of ocean freshening on shelf phytoplankton dynamics.  Geophysical Research Letters, 34, L24607, doi:10.1029/2007GL032010. PDF

Ji, R., P. J. S. Franks, 2007. Vertical migration of dinoflagellates: Model analysis of strategies, growth and vertical distribution patterns. Marine Ecology Progress Series, 344:49-61. PDF

Ji, R., C. Chen, P. J. S. Franks, D.W. Townsend, E.G. Durbin, R. C.Beardsley, R.G. Lough, and R.W. Houghton, 2006. Spring bloom and associated lower trophic level food web dynamics on Georges Bank: 1-D and 2-D model studies. Deep-Sea Research II, 53(23-24):2656-2683. PDF

Ji, R., C. Chen, P. J. S. Franks, D.W. Townsend, E.G. Durbin, R. C.Beardsley, R.G. Lough, and R.W. Houghton, 2006. The impact of Scotian Shelf Water “cross-over” on the plankton dynamics on Georges Bank: A 3-D experiment for the 1999spring bloom. Deep-Sea Research II, 53(23-24), 2684-2707. PDF

Chen, C., L. Wang, R. Ji, J. Budd, D. Schwab, D. Beletsky, G. Fahnenstiel, H. Vanderploeg, B. Eadie, J.Cotner, 2003. Impacts of the Suspended Sediment on the Ecosystem in Lake Michigan: A Comparison between the 1998 and 1999 Plume Events. Journal of Geophysical Research, 109,C10S05, doi:10.1029/2002JC001687, 2004. PDF

Ji. R, C. Chen, D. Schwab, D.Beletsky, J. Budd, G. Fahnenstiel, and M. Bundy, 2002. Influence of suspended sediment on the ecosystem in Lake Michigan: a 3-D coupled bio-physical modeling experiment. Ecological Modeling. 152:169-190. PDF

Chen, C., R. Ji,D. Schwab, D. Beletsky, J. Budd, G. Fahnenstiel, and M. Bundy, 2002. A model study of the coupled biological and physical dynamics in Lake Michigan. Ecological Modeling.152:145-168. PDF

Chen,C., R. Ji,L. Zheng, M. Zhu and M. Rawson, 1999. Influences of physical processes on the ecosystem in Jiaozhou Bay: A coupled physical and biological model experiment. Journal of Geophysical Research 104(c12): 29925-29949. PDF


Book Chapter

Rawson, M., C. Chen, R. Ji, M. Zhu, D. Wang, L. Wang, C. Yarish, J. Sullivan, T. Chopin and Q. Carmona, 2002. Understanding the interaction of extractive and fed aquaculture using ecosystem modeling. In: Stickney R. and McVey J. (eds) Responsible Marine Aquaculture, CABI publishing.